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Abstract As the last time period when concentrations were near 400 ppm, the Pliocene Epoch (5.33–2.58 Ma) is a useful paleoclimate target for understanding future climate change. Existing estimates of global warming and climate sensitivity during the Pliocene rely mainly on model simulations. To reconstruct Pliocene climate and incorporate paleoclimate observations, we use data assimilation to blend sea‐surface temperature (SST) proxies with model simulations from the Pliocene Modeling Intercomparison Project 2 and the Community Earth System Models. The resulting reconstruction, “plioDA,” suggests that the mid‐Pliocene (3.25 Ma) was warmer than previously thought (on average 4.1°C warmer than preindustrial, 95% CI = 3.0°C–5.3°C), leading to a higher estimate of climate sensitivity (4.8°C per doubling of , 90% CI = 2.6°C–9.9°C). In agreement with previous work, the tropical Pacific zonal SST gradient during the mid‐Pliocene was moderately reduced (°C, 95% CI = –0.4°C). However, this gradient was more reduced during the early Pliocene (4.75 Ma, °C, 95% CI = –°C), a time period that is also warmer than the mid‐Pliocene (4.8°C above preindustrial, 95% CI = 3.6°C–6.2°C). PlioDA reconstructs a fresh North Pacific and salty North Atlantic, supporting Arctic gateway closure and contradicting the presence of Pacific Deep Water formation. Overall, plioDA updates our view of global and spatial climate change during the Pliocene, as well as raising questions about the state of ocean circulation and the drivers of differences between the early and mid‐Pliocene. 

Abstract Unlike in the high‐latitude North Atlantic, no deep water is formed in the modern subarctic North Pacific. It has previously been suggested that during climate states different from today, this dichotomy did not endure, and the formation of North Pacific Deepwater (NPDW) occurred in the subarctic North Pacific, which supported an active Pacific meridional overturning circulation (PMOC). Here we provide new records of productivity and sedimentary redox conditions from the central subarctic North Pacific spanning the late Miocene to early Pleistocene. These reconstructions indicate greater‐than‐modern and temporally varying North Pacific export production across the interval of ∼2.7–6 Ma. Our time series, combined with previously published data sets and model output for Pliocene North Pacific Ocean dynamics, support the presence of an active PMOC during the Pliocene, and suggest that the characteristics of NPDW formation varied during this warmer interval of Earth's history. This finding of elevated export production at a time of deep water formation presents a conundrum when considering Quaternary North Pacific Ocean dynamics, where subarctic North Pacific productivity declines during intervals when enhanced overturning is posited to occur. We evaluate our data considering the caveats of both (i.e., Pliocene and Quaternary North Pacific circulation) hypotheses, as well as additional mechanisms unrelated to ocean circulation. Because the Pliocene is a possible analogue for near‐future climate, our results and analyses have important ramifications for our understanding of regional and global climate in the coming decades as the planet continues to warm.